out of africa

“million-year-old”-fossil-skulls-from-china-are-far-older—and-not-denisovans

“Million-year-old” fossil skulls from China are far older—and not Denisovans


careful with that, it’s an antique

The revised age may help make sense of 2-million-year-old stone tools elsewhere in China.

Two skulls from Yunxian, in northern China, aren’t ancestors of Denisovans after all; they’re actually the oldest known Homo erectus fossils in eastern Asia.

A recent study has re-dated the skulls to about 1.77 million years old, which makes them the oldest hominin remains found so far in East Asia. Their age means that Homo erectus (an extinct common ancestor of our species, Neanderthals, and Denisovans) must have spread across the continent much earlier and much faster than we’d previously given them credit for. It also sheds new light on who was making stone tools at some even older archaeological sites in China.

Homo erectus spread like wildfire

Yunxian is an important—and occasionally contentious—archaeological site on the banks of central China’s Han River. Along with hundreds of stone tools and animal bones, the layers of river sediment have yielded three nearly complete hominin skulls (only two of which have been described in a publication so far). Shantou University paleoanthropologist Hua Tu and his colleagues measured the ratio of two isotopes, aluminum-26 and beryllium-10, in grains of quartz from the sediment layer that once held the skulls. The results suggest that Homo erectus lived and died along the Han River 1.77 million years ago. That’s just 130,000 years after the species first appeared in Africa.

(Side note: This river has been depositing layers of silt and gravel on the same terraces for at least 2 million years, and that’s just extremely cool.)

The revised date suggests that Homo erectus spread across Asia much more quickly than anthropologists had realized. So far, the oldest hominin bones found anywhere outside Africa are five skulls, along with hundreds of other bones, from Dmanisi Cave in Georgia. The Dmanisi bones are between 1.85 million and 1.77 million years old, and they (probably—more on that below) also belong to Homo erectus.

Until recently, the next-oldest Homo erectus fossils outside Africa were the 1.63-million-year-old fossils from another Chinese site, Gongwangling, a short distance north of Yunxian. (That’s not counting a couple of teeth from a site in southern China with an age that is a little less certain.) Those dates had suggested Homo erectus seemed to have taken a leisurely 140,000 years to spread east into Asia. But it now looks like hominins were living in Georgia and central China at about the same time, which means they spread out very fast, started earlier than we knew, or both.

The Homo longi and short of it

All of this means that the Yunxian skulls are probably not—as a September 2025 study claimed—close ancestors of the enigmatic Denisovans. The authors of that paper had digitally reconstructed one of the skulls and concluded that it looked a lot like a 146,000-year-old skull from Harbin, China (which a recent DNA study identified as a Denisovan, also known as Homo longi).

The researchers had argued that the original owners of the Yunxian skulls had lived not long after the Denisovan/Homo longi branch of the hominin family tree split off from ours—in other words, that the Yunxian skulls weren’t mere Homo erectus but early Homo longi, close cousins of our own species. Using the original paleomagnetic dates for the Yunxian skulls, that study’s authors drew up a hominin family tree in which our species and Denisovans are more closely related to each other than either is to Neanderthals—one in which the branching happened much earlier than DNA evidence suggests.

There were many issues with those arguments, but the revised age for the Yunxian skulls sounds like a death knell for them. “1.77 million years is just too old to be a credible connection to the Denisovan group, which DNA tells us got started after around 700,000 years ago,” University of Wisconsin paleoanthropologist John Hawks, who was not involved in the study, told Ars in an email.

But the most interesting thing about these skulls being 1.77 million years old is that the date provides a reference point for understanding even older sites in China—sites that may suggest that Homo erectus wasn’t even the first hominin to make it this far.

Photograph of stone tools

Stone tools collected from Shangchen, China.

Credit: Prof. Zhaoyu Zhu

Stone tools collected from Shangchen, China. Credit: Prof. Zhaoyu Zhu

Out of Africa: The prequel

Homo erectus first shows up in the fossil record around 1.9 million years ago in Africa, where it’s sometimes also called Homo ergaster because paleoanthropologists seem to enjoy naming things and then arguing about those names for several decades. A few hundred thousand years later, Homo erectus showed up everywhere: from South Africa northward to the Levant and from Dmanisi Cave in Georgia eastward to the islands of Indonesia.

We typically think of Homo erectus as the first of our hominin ancestors to expand beyond Africa, along routes that our own species would retread 1.5 million years later. More to the point, many paleoanthropologists think of them as the first hominin that could have adapted to so many different environments, each with its own challenges, along the way.

But we may need to give earlier members of our genus, like Homo habilis, a little more credit because stone tools from two other sites in China seem to be older than Homo erectus. At Shangchen, a site on the southern edge of China’s Loess Plateau, archaeologists unearthed stone tools from a 2.1-million-year-old layer of sediment. And at the Xihoudu site in northern China, stone tools date to 2.43 million years ago.

“If you have a site in China that’s 2.43 million years, and the origin of Homo erectus is 1.9 million years ago, either you need to push the origin of Homo erectus back to 2.5 or 2.6 million years or we need to accept that we need to be looking at other hominins that may have actually moved out of Africa,” University of Hawai’i at Manoa paleoanthropologist Christopher Bae, a coauthor of the new study, told Ars.

So who made those 2-million-year-old tools?

Archaeologists have unearthed stone tools but no hominin fossils at both sites, making it difficult to say for sure who the toolmakers were. But if they weren’t Homo erectus, the next most likely suspects would be older members of our genus, like Homo habilis or Homo rudolfensis. That would mean hominin expansion “out of Africa” actually happened several times during the history of our genus: once with early Homo, again with Homo erectus, and yet again with our species.

“There could have been an earlier wave that died out or interbred, so there’s all kinds of possibilities open there,” Purdue University paleoanthropologist Darryl Granger, also a coauthor of the recent study, told Ars.

In fact, there’s some debate about whether the Dmanisi fossils actually belonged to Homo erectus proper. One thing the two dueling reconstructions of the Yunxian skulls agree on is that those hominins had flattish faces, more like ours—and like the 1.63-million-year-old Homo erectus skull from Gongwangling. But the Dmanisi hominins’ lower faces project dramatically forward, like those of older hominins.

Some paleoanthropologists classify the Dmanisi fossils as their own species, but others argue they’re more like early members of our genus, such as Homo habilis or Homo rudolfensis. Those earlier hominins may have been more capable of migrating and adapting than we’ve realized.

It’s still very clear, from both fossil and genetic evidence, that our species evolved in Africa and spread from there to the rest of the world. But it’s also increasingly clear that there were several other species of hominins in other places, doing other things, at least off and on, for a very long time before we showed up. Yunxian, and its revised age, could help anthropologists better understand part of that story.

“Actually being able to anchor the Homo erectus sites with firm, solid dates helps us try to reconfigure this model,” said Bae. “This is where Yunxian really plays a major role in this. Now that we’ve got older dates to anchor the Yunxian Homo erectus fossils, I think we can really bring in this discussion with Xihoudu and Shangchen.”

Time to dig deeper

The answers may still lie buried—maybe just a few meters below the fossil skulls and stone tools at sites like Yunxian and Gongwangling, in older sediment layers. Archaeologists may not have seen a reason to explore these, since no one lived in China before 1.7 million years ago. The age of the Yunxian skulls, along with the even older stone tools at Shangchen and Xihoudu, may warrant deeper digging.

“People haven’t been looking for artifacts and fossils in two-plus million-year-old sediments in these locations in China,” said Granger. “I can think of places that I would like to go back and look if I had more time and money.”

At other sites, researchers have already unearthed fossil animal bones from the same age range as China’s oldest stone tools, but paleoanthropologists haven’t double-checked whether any of those bones might belong to early hominins rather than other mammals. Bae said, “It’s just that they haven’t been receiving any attention, or not enough attention.”

Science Advances, 2026. DOI: 10.1126/sciadv.ady2270 About DOIs).

Photo of Kiona N. Smith

Kiona is a freelance science journalist and resident archaeology nerd at Ars Technica.

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studies-pin-down-exactly-when-humans-and-neanderthals-swapped-dna

Studies pin down exactly when humans and Neanderthals swapped DNA


We may owe our tiny sliver of Neanderthal DNA to just a couple of hundred Neanderthals.

The artist’s illustration shows what the six people buried at the Ranis site, who lived between 49, 500 and 41,000 years ago, may have looked like. Two of these people are mother and daughter, and the mother is a distant cousin (or perhaps a great-great-grandparent or great-great-grandchild) to a woman whose skull was found 130 kilometers away in what’s now Czechia. Credit: Sumer et al. 2024

Two recent studies suggest that the gene flow (as the young people call it these days) between Neanderthals and our species happened during a short period sometime between 50,000 and 43,500 years ago. The studies, which share several co-authors, suggest that our torrid history with Neanderthals may have been shorter than we thought.

Pinpointing exactly when Neanderthals met H. sapiens  

Max Planck Institute of Evolutionary Anthropology scientist Leonardo Iasi and his colleagues examined the genomes of 59 people who lived in Europe between 45,000 and 2,200 years ago, plus those of 275 modern people whose ancestors hailed from all over the world. The researchers cataloged the segments of Neanderthal DNA in each person’s genome, then compared them to see where those segments appeared and how that changed over time and distance. This revealed how Neanderthal ancestry got passed around as people spread around the world and provided an estimate of when it all started.

“We tried to compare where in the genomes these [Neanderthal segments] occur and if the positions are shared among individuals or if there are many unique segments that you find [in people from different places],” said University of California Berkeley geneticist Priya Moorjani in a recent press conference. “We find the majority of the segments are shared, and that would be consistent with the fact that there was a single gene flow event.”

That event wasn’t quite a one-night stand; in this case, a “gene flow event” is a period of centuries or millennia when Neanderthals and Homo sapiens must have been in close contact (obviously very close, in some cases). Iasi and his colleagues’ results suggest that happened between 50,500 and 43,000 years ago. But it’s quite different from our history with another closely related hominin species, the now-extinct Denisovans, with whom different Homo sapiens groups met and mingled at least twice on our way to taking over the world.

In a second study, Arev Sümer (also of the Max Planck Institute) and her colleagues found something very similar in the genomes of people who lived 49,500 to 41,000 years ago in what’s now the area around Ranis, Germany. The Ranis population, based on how their genomes compare to other ancient and modern people, seem to have been part of one of the first groups to split off from the wave of humans who migrated out of Africa, through the Levant, and into Eurasia sometime around 50,000 years ago. They carried with them traces of what their ancestors had gotten up to during that journey: about 2.9 percent of their genomes were made up of segments of Neanderthal ancestry.

Based on how long the Ranis people’s segments of Neanderthal DNA were (longer chunks of Neanderthal ancestry tend to point to more recent mixing), the interspecies mingling happened about 80 generations, or about 2,300 years, before the Ranis people lived and died. That’s about 49,000 to 45,000 years ago. The dates from both studies line up well with each other and with archaeological evidence that points to when Neanderthal and Homo sapiens cultures overlapped in parts of Europe and Asia.

What’s still not clear is whether that period of contact lasted the full 5,000 to 7,000 years, or if, as Johannes Krause (also of the Max Planck Institute) suggests, it was only a few centuries—1,500 years at the most—that fell somewhere within that range of dates.

Artist’s depiction of a Neanderthal.

Natural selection worked fast on our borrowed Neanderthal DNA

Once those first Homo sapiens in Eurasia had acquired their souvenir Neanderthal genes (forget stealing a partner’s hoodie; just take some useful segments of their genome), natural selection got to work on them very quickly, discarding some and passing along others, so that by about 100 generations after the “event,” the pattern of Neanderthal DNA segments in people’s genomes looked a lot like it does today.

Iasi and his colleagues looked through their catalog of genomes for sections that contained more (or less) Neanderthal ancestry than you’d expect to find by random chance—a pattern that suggests that natural selection has been at work on those segments. Some of the segments that tended to include more Neanderthal gene variants included areas related to skin pigmentation, the immune response, and metabolism. And that makes perfect sense, according to Iasi.

“Neanderthals had lived in Europe, or outside of Africa, for thousands of years already, so they were probably adapted to their environment, climate, and pathogens,” said Iasi during the press conference. Homo sapiens were facing selective pressure to adapt to the same challenges, so genes that gave them an advantage would have been more likely to get passed along, while unhelpful ones would have been quick to get weeded out.

The most interesting questions remain unanswered

The Neanderthal DNA that many people carry today, the researchers argue, is a legacy from just 100 or 200 Neanderthals.

“The effective population size of modern humans outside Africa was about 5,000,” said Krause in the press conference. “And we have a ratio of about 50 to 1 in terms of admixture [meaning that Neanderthal segments account for about 2 percent of modern genomes in people who aren’t of African ancestry], so we have to say it was about 100 to maybe 200 Neanderthals roughly that mixed into the population.” Assuming Krause is right about that and about how long the two species stayed in contact, a Homo sapiens/Neanderthal pairing would have happened every few years.

So we know that Neanderthals and members of our species lived in close proximity and occasionally produced children for at least several centuries, but no artifacts, bones, or ancient DNA have yet revealed much of what that time, or that relationship, was actually like for either group of people.

The snippets of Neanderthal ancestry left in many modern genomes, and those of people who lived tens of thousands of years ago, don’t offer any hints about whether that handful of Neanderthal ancestors were mostly male or mostly female, which is something that could shed light on the cultural rules around such pairings. And nothing archaeologists have unearthed so far can tell us whether those pairings were consensual, whether they were long-term relationships or hasty flings, or whether they involved social relationships recognized by one (or both) groups. We may never have answers to those questions.

And where did it all happen? Archaeologists haven’t yet found a cave wall inscribed with “Og heart Grag,” but based on the timing, Neanderthals and Homo sapiens probably met and lived alongside each other for at least a few centuries, somewhere in “the Near East,” which includes parts of North Africa, the Levant, what’s now Turkey, and what was once Mesopotamia. That’s one of the key routes that people would have followed as they migrated from Africa into Europe and Asia, and the timing lines up with when we know that both Homo sapiens and Neanderthals were in the area.

“This [same] genetic admixture also appears in East Asia and Australia and the Americas and Europe,” said Krause. “If it would have happened in Europe or somewhere else, then the distribution would probably look different than what we see.”

Science, 2023 DOI: 10.1126/science.adq3010;

Nature, 2023 DOI: 10.1038/s41586-024-08420-x;

(About DOIs).

Photo of Kiona N. Smith

Kiona is a freelance science journalist and resident archaeology nerd at Ars Technica.

Studies pin down exactly when humans and Neanderthals swapped DNA Read More »